Flash is the smaller, cheaper, higher-concurrency worker. Pro is the larger reasoning and review route. Both share the V4 family and API feature rows, but they should be assigned by job stage.
Concrete split: Flash is listed at 284B total and 13B activated parameters; Pro is listed at 1.6T total and 49B activated parameters. DeepSeek lists lower Flash prices and higher Flash concurrency. The current API page says the Responses API supports Flash now and Pro support is planned; the July 31 changelog says the Pro API is unchanged and the official V4 Pro release will follow soon.
Use Pro for planning, contradictions, long-context review, and final judgment. Use Flash for extraction, classification, draft cleanup, issue grouping, and other bounded loops. If the handoff point is unclear, keep the route on Pro or human review.